Jūgo

, was being held at Orochimaru's North Base, which was also where most of Orochimaru's secret research was conducted. He was recruited as a member of Taka.

Background
Jugo's personality is usually quiet and unassuming, but, when angered, those around him stand little chance of survival. One time, he even became so angry that he devoured an entire village. This has caused others to label him as the "Scales", where anything could tip the balance. Cursing his existence, he took to secluding himself in caves, hoping to bring no-one any harm. This lasted until Kimimaro came to him, who was himself the last remnant of his own Kaguya clan. Following Kimimaro, Jugo allowed himself to be one of Orochimaru's test subjects. Upon examination, Orochimaru was able to isolate enzymes in Jugo's blood that caused him to transform. By extracting these enzymes, Orochimaru was able to use them to produce similar, but more controlled, effects in other people, leading to the creation of the cursed seal. As time passed, Kimimaro died, then Orochimaru followed him, and Jugo shut himself away once more.

Appearance
Jugo has a bracket-shaped symbol on both of his palms. Whether or not this has any significance has yet to be seen. He is the tallest, non-animal character in the series who's precise height (202 cm) is known. Jugo wears a black shirt, tan pants, and a blue cloak. After Taka had allied with Akatsuki, Jugo was shown wearing an Akatsuki robe.

Jugo has spiky orange hair, and an usually calm face. However, when affected by his murderous side, his skin turns dark, and he grows various weapons and appendages to suit his battle combat situation, as well as having a deranged look.

Personality
While Jugo initially appeared quite bloodthirsty, Karin noted that this was only common behavior when he was enraged; Jugo himself didn't want to kill anyone. Jugo's fits of rage are random, and thus, the reason why he willingly remains confined. The only reason he agreed to go with Sasuke is because he views Sasuke as the only person capable of preventing him from killing people while enraged, by the means of killer intent or the Sharingan. Before Sasuke, it was Kimimaro who kept Jugo in check, and as Kimimaro was the only person Jugo came into any contact with, the two grew close. After hearing that Kimimaro sacrificed his life to deliver Sasuke to Orochimaru, Jugo believes that Kimimaro's spirit continues to live on in Sasuke.

Suigetsu can't stand Jugo, and commented that one of the reasons he had to retrieve the Decapitating Carving Knife was because of him. Karin noted that Jugo is mentally unstable, a view confirmed when Jugo was muttering to himself over who he would kill next, which he would determine by gender. While discussing Sasuke Uchiha's team with the other members of Akatsuki, Pain referred to Jugo as, referencing his dual personality. The English manga used the term "Bipolar Jugo", obtaining the same effect.

Hunt for Itachi arc
After defeating Orochimaru, and recruiting Suigetsu and Karin, Sasuke Uchiha travelled to Orochimaru's North Base, to recruit Jugo. Initially unwilling to join forces with Sasuke, Jugo tapped into his Cursed Seal, but was then frightened into releasing it by Sasuke's killer intent. He then joined the newly formed Hebi in their mission to hunt down and kill Akatsuki's Itachi Uchiha, Sasuke's older brother.

Jugo would later discover multiple hidden bases belonging to the Akatsuki, where Sasuke confronted a clone of Itachi in one of them, who challenged Sasuke to meet him at the Uchiha Clan Stronghold. Jugo remained behind with the other members of Hebi and Kisame Hoshigaki as Sasuke confronted Itachi.

Hunt for the Eight-Tails arc
After Itachi's death, and Sasuke's "education" in his brother's true past by Madara Uchiha, Jugo and the other members of Hebi were reformed into Taka. Having allied themselves with the Akatsuki, Taka was sent to capture the Eight-Tailed Beast, so that it could be extracted from its host, Killer Bee. Infiltrating Kumogakure, they confronted Killer Bee, only to have him defeat Suigetsu. When Jugo attempted to defeat him as well, Killer Bee overpowered him using the sword Suigetsu had wielded, which originally belonged to Zabuza Momochi. After Sasuke was critically injured, Jugo tells Karin that she did not have enough chakra to heal him and starts to infuse some of his flesh with Sasuke, to heal his wounds, saying it would work as he was compatible for his cursed seal. This however somehow reduced him to a younger appearance. When their team started working together, Jugo concentrated on removing his teammates from danger, even though he initially objected to rescuing Karin, as he thought she would die from Sasuke's Amaterasu. Sasuke was eventually able to capture Killer Bee, albeit a fake, and Jugo recuperated with the rest of Taka.

Five Kage Summit arc
As Taka rested in one of Akatsuki's hideout, Karin sensed a spy had followed them. Jugo and Sasuke went and intercepted him, and left the hideout for a new one shortly after. Taka later set out for Konoha. On their way they were intercepted by Tobi who informed them that Konoha had been destroyed and later it was explained by Zetsu, who suddenly appeared, the circumstances of the destruction and that a man named Danzo had been named Hokage. As per Tobi's advice and orders Sasuke changes their goal to the Kage Summit and Team Taka is then guided by Zetsu, who had split up into two bodies.

Jugo and the rest of Team Taka arrived at the Land of Iron, and after Suigetsu scouted the area and reported there are many guards, Sasuke tells Jugo to use animals to find the least guarded route. Their presence is soon discovered and Sasuke engages the Samurai that come after them. While Sasuke deals with them, Karin continues to worry about the Samurais only to be told by Jugo to find Danzo as he was their primary target. When Karin once again voices her concerns, Jugo tells her that he will deal with them.

Taka is attacked by Kumogakure ninja that Jugo identifies as two jonin and the Raikage. After being blinded by a genjutsu and Sasuke dispelling it with his Sharingan, the Raikage attacks. Suigetsu comes and takes the hit at the cost of his sword breaking. Realizing that one of them was a sensor type like Karin, making it difficult for her locate Danzo and also keep herself hidden, Jugo resolves that killing him is the only options and says that he will do it. Jugo then transforms into his Curse Seal level 2.

He is told by the Raikage to stop being cocky but goes into his murderous mental state. Suigetsu then questions Sasuke, that since he is now in "killer mode", would he be able to tell the difference between friend and foe. He is then overwhelmed by the Raikage's speed and strength and forced into the wall and the Raikage even punches through a defensive shield on his arm and further into his chest. With a hole in his arm and chest, he has seemingly been killed, but Jugo only smirks and aims dozens of boosters from his back straight at the Raikage. Laughing, Jugo calls for the Raikage to die and then fires a huge blast at the Raikage at point blank range. Believing the Raikage to be dead, Jugo begins laughing once more and asks who is next and prepares to attack someone else, but he is then hit again by the Raikage with a sudden elbow strike. Jugo smashes against the wall and seems to be knocked unconscious. He later woke up to see Sasuke using Susanoo, and was wandering if that's what Sasuke was referring to wanting to try out earlier. After Sasuke uses Susanoo to collapse the cave, Jugo is left seriously injured, but conscious. Having fallen near the body of a Samurai, Jugo states that he might make it, and extends part of his body to pierce the deceased Samurai and seemingly absorb his insides. Jugo's health, as well as his adult stature, are restored by this technique. He is later seen with Suigetsu, who is suggesting that they should follow the Raikage, Kazekage and Sasuke. Jugo tells him not to do so because they would be killed. However, Suigetsu says that he has a plan in mind. Suigetsu planned for him and Jugo to disguise themselves as Samurais though Jugo states it wasn't a good plan when their cover is blown by the other Samurai.

Cursed Seal
Jugo is the very origin of the Orochimaru's Cursed Seals, so he is able to use its powers to its full extent without it eroding his body. Also, unlike the cursed seals distributed to other people, Jugo can have many more forms to change to, as opposed to being restricted into one. He also refers to people with a Curse Seal as a copy of him.

His brief fight with Sasuke and Suigetsu revealed that Jugo can control his transformation, unlike the static effects of a standard Cursed Seal. When battling Sasuke, he changed his left arm into a piston-like ram to increase his punching power, and then altered it into a massive axe blade to counter Suigetsu's sword. He is also capable of doing only a partial transformation, invoking a level two form over one portion of his body without completely changing.

Jugo's calmer personality is able to throw people by using his arm and releasing chakra out the other direction. This can also be used for escaping, charging right out of the battle with great speed.

He can also fuse part of his body, while in Cursed Seal form, to replace parts of other Cursed Seal-infected people's bodies, as he does to Sasuke. However, the downside to this appears to be it reverts Jugo to the appearance of a child. He can also take bodies of others to fuse into his own body to repair injuries.

By entering into his Level 2 state, Jugo gains massive power boost, but loses his mind into his murderous side. As with all others with a Cursed Seal, Jugo's skin turns dark, with black sclera in the eyes. His elbows have sharper edgings, his nails sharpen, he grows horns at the top of his head that extends to his cheeks, and on his back are what appears to be six jet boosters like appendages, similar to the bones on the back produced by Kimimaro in his Level 2 form, that release fire. These appendages can increase in number and extend forward to aim at a target of Jugo's choosing, then launch massive blasts of chakra with cannon-like force, devastating an opponent, especially at point-blank range. In his Level 2 form, Jugo is also very resilient, as he took a punch from the Raikage right through the arm and chest and managed to retaliate with ease.

Communications with Animals
Jugo is able to understand and communicate with animals, such as birds, and is able to request them to help him accomplish tasks, such as acting as scouts.

Trivia

 * Jugo's opinion seems to be the only one out of Taka that Sasuke listens to. While Sasuke usually brushes everyone off, he listens to Jugo's advice.
 * Jugo will be a playable character in Naruto Shippūden: Narutimate Accel 3 and Naruto Shippuden: Ryujinki.