Board Thread:Theories and Speculation/@comment-6769189-20200515102419/@comment-4080775-20200521101220

@Actionmanrandell No, you have to be a transmigrant to Awaken it in terms of how Hagoromo understood it, but because the Otsutsuki can awaken it underwise, it does seem to be a genetic thing.

Orochimaru and Kabuto could theoretically gain it by just using Madara or Sasuke's DNA. They don't need to be reincarnations if they have access to their genetics, because you just need the genetics themselves for the Strange Transmission Distant Shadow to work.

There's absolutely no reason to assume he doesn't still have the same body. We know he wouldn't give up such a body without a reason, especially since he grants him Hashi's abilities. Even if he did give it up, he'd likely still have access to the abilities because the soul of the Zetsu would go with him as per the Living Body Reincarnation. Even if it didn't, he'd just use Jugo's Cellular Regeneration Absorption to absorb what he needs from the zetsu body. This would be the same way he was revived from Kabuto's flesh, giving him Hashi Cells and abilities once more.